Grudge
A soul will only inhabit a root if it's holding a grudge. Have you ever seen a nice, haunted tree? No! They're always angry and trying to get someone back for something.
Don't die with a grudge, or your soul might end up angry and trapped in some roots.
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If you see one of these grudge trees, it's probably inhabited by the soul of a condor (こん). Humans tend to kill condors and take their trees away to build houses and make toilet paper. So condors hold pretty bad grudges against us, and so their souls end up stuck in a lot of trees.
Picture a tree with the soul of a condor with a grudge in it before you. You feel bad about being so wasteful with paper and chopsticks now, huh?